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    The Improbable Primate

    Page 20
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      core xii, 55, 67, 133, 145–6

      Middle Pleistocene aridity 69–70

      northern xi–xii, 64–6, 76–7, 78, 96, 106

      northern glaciations 110

      overlap of rain chaser/Neanderthal populations 105–6

      as site of human origin 55, 147

      southern xi, 77–8, 83–4, 90–4, 106, 108–11, 112

      Middle East 105, 109

      aridification 70, 90

      erectus 55, 62–3

      Natufians 147, 149

      Neanderthal extinctions 97–8

      Milton, Katherine 1

      Miocene, Middle 1–10

      emergence of C4 grasses 17

      forest primates 1–4, 6–8, 10–11

      modernity xv–xvi molluscs 60, 63, 101–2

      monitor lizards 16, 60

      monk seals 101, 148

      Morgan, Elaine 13

      Mumbwa Caves, central Africa 94

      Murray-Darling Basin, Australia 142–4, 147

      population containment 149

      Nariokotome Boy 41, 43–4

      Natufian people, Middle East 147, 149

      natural selection 37, 47, 48, 52, 84, 85

      Nature 41

      Neanderthals (Homo sapiens neanderthalis) xi, 81, 85, 87, 90, 95–107, 109, 112, 123–4, 129, 151

      altitude movement with climate change 119

      coastal sites 92, 100–2

      commitment to small-scale territories 106–7

      considered separate species ix

      contact with Denisovans 117

      contact with rain chasers 105–6

      extinction patterns 97–8

      factors important for survival 99–102

      genetic legacy ix, 81, 111, 121

      genetic mixing of populations 102

      geographical range 96–7

      habitat 102–4, 121–2

      hind limb anatomy 106

      last populations 97, 98–9, 107, 121–3

      as lineage of Homo sapiens x, xiv, xviii

      mountain launchpad into Eurasia 116–20

      muscular build xii–xiv, 106–7

      sedentary lifestyle 148–9

      seen as cognitively inferior xvi

      split from Homo sapiens sapiens 81

      spread into Europe via central Asia 112

      use of wooden implements xv

      Neanderthals and Modern Humans (Finlayson) 78

      New Guinea Highlands 151

      Niah Cave, Borneo 124–5, 133, 151

      nomadic lifestyles:

      Australian dessert 138–44, 146

      and geographical expansion 150

      and reciprocity 144, 146

      North Africa 67

      aridification and climate fluctuation 70, 77, 90–1

      erectus 63–4

      Middle Pleistocene 77

      rain chasers 87, 90–2, 93

      North America 130, 147–8

      Nubians 90–1

      Ohalo, Sea of Galilee 147

      Oldowan technology 52, 63, 64–5, 66, 67, 70, 71, 71 Fig. 4, 151, 170n

      Olorgesailie, Kenyan Rift Valley 72–3

      open treeless grasslands 33–4, 35

      orang-utans (Pongo) 25

      ostriches 67

      otters 15

      Oued Djebanna, Algeria 91

      Out-of-Africa/Out-of-Africa 2 models x, xi, 55, 108–10, 134

      Pacific coast of America 147–8

      Pakefield, Suffolk 76

      Panxian Dadong, China 77–8

      Paranthropus 29, 33–4, 35, 42

      diet 34

      distribution 33

      extinction 34

      habitat 33–4

      height 44

      moving towards life on the ground 37

      rocky places 37

      start of line 36

      Paranthropus bosei 34

      Paranthropus robustus 34

      Pavlov, Czech Republic 147

      Persian Gulf 55

      Peru 150

      Petraglia, Mike 124

      phytoliths 31, 162n

      pigs 16, 46, 63

      Pinnacle Point, South Africa 92, 102

      Pleistocene 130, 142

      Late 130

      Middle xvii, 69–82

      Pliocene 10, 130

      emergence of C4 grasses 17

      population genetics xi

      predators 50–1

      attraction of seasonally-flooded areas 21

      patchily distributed resources and movement in groups 4–7

      primates 52

      behaviour around water 24–5

      bonding with natal area 146

      brain expansion 3–4

      brain function in forest contexts 1–4

      coming down to the ground 8–9

      effects of food density on behaviour 6–8

      group living 6–8, 12–13

      Miocene forest 1–4, 6–8, 10–11, 146

      predisposition for culture 12–13

      protein supplements for fruit-based diets 3, 12–13, 40

      transition from tropical rainforest to tropical seasonal forest 21–2

      proboscis monkeys (Nasalis) 24

      Puritjarra Rock Shelter, Australia 134–5, 145, 147

      Pyrenees 5–6, 106

      pythons 15

      Radcliffe-Brown, Alfred 142

      rain chasers (southern Middle Earth lineage) 83–95, 96, 105, 106–7

      ‘Africa viewpoint’ 108–9

      brain size 84, 85, 94

      cultural and genetic interchange with Neanderthals 81, 90, 105, 121–2

      as desert dwellers 132–6, 144

      as endurance runner and long-distance walker 83, 85, 111, 122

      Eurasian open habitat 122–3

      expansion into Australia 111, 122–3, 125, 133–6

      expansion into Eurasia 120–1

      expansion into south-east Asia 111–12, 123–5, 135

      global expansion 108–33

      hunting mobile prey 89, 121

      lightweight build 83, 85, 94, 105, 106–7, 122

      mobility 94, 105

      overlapping with Neanderthals 105–6

      physical appearance 85–6

      population isolation and re-contact 86, 110

      southern sites 130

      split from Neanderthals 81

      supposed arrival time 200 thousand years ago 86

      technological innovation 87–90, 94, 105, 121, 122

      rain front tracking 87–8, 137

      rainforests 144

      as barriers 111

      expansion 61, 68, 115–16, 125

      human alteration, Niah Cave 124–5, 133

      Miocene 1–4, 6–8, 10–11, 59

      shrinking/opening up 21–2, 43, 60, 80, 111, 130

      Ramidus (Ardipithecus ramidus) 32, 37, 59

      diet 17, 25, 157n

      facultative bipedalism 17, 23–4, 160n

      habitat 17–19, 21, 22–5

      opportunism 25–6

      reciprocity and sharing 144, 146

      Reed, K. 33–4

      reindeer xv, 121, 147

      rhinos 63, 65, 66, 67, 91

      rocky places 37–9, 72, 77, 82, 135, 142

      erectus 62, 75

      Neanderthals and 103, 104, 122, 148

      rain chasers 92–3

      in works of art 152

      Roebroeks, W. 55, 67

      sabre-tooth cats 16, 66

      Sahara Desert 93, 110, 130, 135

      alternating wet-dry conditions 79

      expansion 79, 90–1 Sahelanthropus 37, 59, 79

      salmon 147–8

      savannah/aquatic dichotomy 26

      savannah/grassland 13, 31, 94, 143

      corridors 112

      open 9, 26, 34, 36, 60–1, 67, 77

      wooded 15–16, 17, 22, 36, 37, 64

      Savannahstan 55, 67

      Schöningen, Germany xv

      Science xviii, 16

      sea crossings 61–2, 68, 125, 133

      seals 92, 101, 148

      seasonal flooding 18–21, 25

      sedentary lifestyles 147–9

      and geographi
    cal expansion 150

      and resource predictability 147–9

      Sel’ungur, Kyrgyzstan 77

      Serpent’s Glen, Australia 134

      sexual dimorphism 17, 44

      shellfish 91, 93, 101

      Siberia 96, 106, 112, 119

      Sima de Elephante, Spain 64–5

      sociability, Mardu peoples 141

      sources and sinks model 78, 112, 128–9

      South Africa 33, 35, 67

      aridification 70

      cave dwelling 92–3

      coastal site exploitation 92, 101, 102

      spread of Middle Pleistocene humans with Acheulian technology 70–2

      south-East Asia xii, 41, 59–62, 77, 111, 133

      altitude movement in response to water shortage 77–8, 113

      coastal site exploitation 75, 92

      Denisovans 112, 115–18, 123

      effects of cooling and drying 60–1, 77–8

      effects of warming 61

      erectus xii, 41, 54, 59–62, 80

      impact of Toba Volcano 124

      Middle Pleistocene 75, 77–8, 80

      rain chaser expansion into 111, 123–5

      rainforest barriers 111

      rainforest expansion 61, 68

      as source (pump) area 78, 111–17

      Trinil site, Java 59–61

      very early colonization 111

      Spain see Iberian Peninsula

      spear-throwers, Mardu 139

      spider monkeys 3, 154n

      Stewart, J. R. 110

      stone tools 87

      earliest known 39–40

      Flores 61

      Neanderthal 148

      typological classification drawbacks xiv

      see also Acheulian technology; Oldowan technology

      storks 14, 60

      Strait of Gibraltar 4–5, 64, 68, 99

      Stringer, C. B. 110

      swamp monkeys (Allenopithecus) 24

      Syria 118

      Systematics and the Origin of Species from the Viewpoint of a Zoologist (Mayr) vii

      Taforalt, Morocco 91

      territoriality 142, 146–7

      and genetic isolation 143–4

      Neanderthals 149

      and violence 143, 144, 146, 149

      tigers 60

      tits 4

      Toba Volcano eruption 124, 133

      Tonkinson, Robert 136

      tools:

      composite technology 89–90, 105, 109, 121

      hafted 88, 89, 106

      lightweight and portable 87–8, 94, 120

      Mardu toolkit xiv–xv, 139–40

      projectile technology 120, 122

      small 93

      see also Acheulian technology; Oldowan technology; stone tools

      tortoises 16

      Toumaï (Sahelanthropus tchadensis) 13–16

      diet 25

      habitat 13–16, 21, 22–5

      opportunism 25–6

      trading networks 92

      tree sloths 3

      Trinil, Java 59–61

      Turkey 98, 118

      turtles 15, 19, 42, 60, 63

      Ubeidiya 63, 151

      UNESCO habitat definitions 22

      Vanguard Cave, Gibraltar 101

      visual discrimination, in forest canopy 2

      vultures 5 Fig. 2, 5–6, 50, 51

      water

      and beginnings of agriculture 151

      centrality to Mardu people 137

      and choice of Acheulian/Oldowan technology 64–5, 70–2

      in Central Australia 134

      coastal areas as source of fresh 93, 100

      as evolutionary driver xii–xiv, xvii, 37, 47, 49, 52, 58, 59, 84–5, 125–31

      importance to hominids and early humans 29, 32, 36–8, 79, 82, 93–4, 103

      inbuilt knowledge about sources 58, 85

      links to Himalayan launchpad model 113–118

      Mardu maximization of resources 137, 140–1

      and Neanderthal survival 100, 103

      necessity to human sites 125–30

      Paranthropus habitat 33, 34

      and population density, Australia 135–6

      primate behaviour around 24–5

      predictability of supply, and sedentary lifestyle 149

      reliability and population expansion 149–50

      scarcity and seasonality 43

      seasonal flooding 14–15, 19–21

      and territorial vs. nomadic lifestyles 146, 149

      water buffalo 91

      water shortage 36–7, 84–5

      and altitude movements 78, 115–18

      and Acheulian technology 58–9, 70–1, 87–8, 89

      Barbary macaque response 27–8

      human adaptation to extreme 142

      impact on Neanderthals 103, 104–5

      and increased mobility 37, 38, 47, 49

      increasing after 400 thousand years ago 83–4

      and isolation of human populations 79–80

      lightness of build advantages 83–4

      Middle Pleistocene 77–8

      and Neanderthal extinction, Middle East 98

      seasonal, as consequence of climate change 36–7, 43, 47

      Water Optimization Hypothesis 125–30

      water/trees/open spaces 22–5, 23 Fig. 3, 38, 42, 42–3

      Australia 135, 142

      australopithecines 29–30, 33–4, 35, 36

      changes in distribution 72–3

      early hominid habitats 22–5

      erectus 60–1, 62, 63–4, 72–3, 75, 77

      in landscape paintings 152

      habitat management to produce 124–5, 133, 151–2

      Neanderthal habitat 103–4, 106, 121–2

      Puritjarra 135

      re-creation in parks 152

      water/treeless open spaces 122–3

      weather prediction 140

      wetlands 15, 18–19, 34, 38, 62–3, 66–7, 75, 143

      coastal 61, 68

      permanent vs. seasonal 64–6

      productivity 21–2

      reduction 36

      White, Tim 17–18, 19, 22, 23–4

      wild boar 25

      as scavengers 20–1

      Wolpoff, Milford ix, x, 86–7

      wood, working of 73

      wooden tools xiv–xv, 39

      woolly mammoths xv

      Wrangham, Richard 74

      Yuanmou Basin, China 67

      Zagros Mountains, Iran 106, 118, 119, 120, 123

     

     

     



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